Characteristics of bundle sheath cells in $\text{C}_4$ plants}
(A) Have thin cell walls
(B) Contain centrifugally arranged chloroplasts
(C) Contain stacked thylakoid membranes and do not have starch granules
(D) Carry out light-independent reactions of photosynthesis
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To minimize photorespiration, bundle sheath cells perform the Calvin cycle under elevated $\text{CO}_2$ levels. Their chloroplasts are agranal (lack grana to avoid producing oxygen) but accumulate abundant starch.