Villous atrophy is associated with
(A) Eimeria ovinoidalis
(B) Trichostrongylus colubriformis
(C) Nematodirus battus
(D) Strongyloides papillosus
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Any major parasite that invades or burrows into the small intestinal mucosa (such as coccidia and small intestinal roundworms) will inevitably cause some degree of villous atrophy as a pathological consequence.